A Checklist of Virginia Lichens

 

Suggested citation:

Hodkinson, B. P., R. C. Harris, and M. A. Case. 2009. A Checklist of Virginia Lichens. Evansia 26(2): 64-88. http://www.duke.edu/~bph8/VirginiaLichens/checklist.html [updated: 05 January 2010].

 

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Brendan P. Hodkinson*

*Corresponding author; Department of Biology, Duke University, Durham, North Carolina 27708; email: bph8@duke.edu

 

Richard C. Harris

Institute of Systematic Botany, New York Botanical Garden, Bronx, New York 10458

 

Martha A. Case

Biology Department, College of William and Mary, Williamsburg, Virginia 23187

 

Abstract. A list of 603 taxa of lichen-forming, lichenicolous, and allied fungi has been compiled exclusively from specimens examined and verified by the authors or associated specialists. This list contains 195 potential state records for Virginia, and three species that have not previously been reported from North America: Arthonia anglica Coppins, Calvitimela talayana (Haugan & Timdal) Andreev, and Sphaerellothecium coniodes (Nyl.) Cl. Roux & Diederich.

 

INTRODUCTION

Although the lichen flora of Virginia as a whole is understudied, scattered reports of lichen-forming, lichenicolous, and allied fungal taxa have appeared in a variety of sources over the years (e.g., Ahti & Hyvönen 1985, Allard 1939, 1940, 1957, Allard & Leonard 1944a, 1944b, Alstrup 1991, Amtoft 2002, Amtoft et al. 2008, Blake 1957, Bowler & Rundel 1974, Brodo 1984, Brodo et al. 2001, Clerc & Herrera-Campos 1997, Culberson 1965a, 1965b, Culberson & Culberson 1968, 1978, Culberson & Hale 1965, Degelius 1974, Dey 1978, Dibben 1980, Eckfeldt 1894, Egan et al. 2005, Ekman 1996, Fahselt 1976, Fink 1935, Fleming & Coulling 2001, Forman & Sierk 1970, Fryday 2008, Gowan 1989, Gronovius 1739, Hafellner 1979, 2001, Hale 1970, 1971, 1972, 1974, 1975, Hale & Kurokawa 1964, Harris 1979, 1992, Harris & Ladd 2008, Heidmarsson 2003, Hinds 1998, Hodkinson 2005, 2010, Hodkinson & Case 2008, Hodkinson & Lendemer in review, Hodkinson & Lutzoni 2009, Imshaug 1951, Jørgensen 2000, Kärnefelt 1979, Knudsen 2006, Kurokawa 2001, Lawrey 1980, 1981, 1984, 1991, 1993, Lawrey & Hale 1981, 1988, Lendemer 2009, Lendemer et al. in press, Lendemer & Hodkinson 2009a, 2009b, in press, Lindblom 1997, 2006, Llano 1950, Luttrell 1954, Patterson 1940, 1941, Schwartzman et al. 1987, Sheard 1974, Slaughter 1977, Thomson 1950, 1963, Timdal 1986, Wetmore 1983, 2004). However, the reliability of some specimen reports cannot always be confirmed, and taxonomic changes have rendered others obsolete or intractable without examination of voucher material. The goal of this project is to compile a verified specimen-based checklist of lichen-forming, lichenicolous, and allied fungi for the state of Virginia. 

 

METHODS

This checklist was compiled primarily from voucher specimens identified or verified by the authors (BPH and RCH). The core collection for the list was comprised of specimens held by the New York Botanical Garden (NY), Duke University (DUKE), the National Museum of Natural History at the Smithsonian (US), and Shenandoah National Park (Shen.; note: all specimens stored at this location were collected within the park). Selected specimens from the available reports cited above and several online databases (e.g., ASU, HERBESS, NPLICHEN, OSC, SBBG, and SMNH) were also examined. On a few rare occasions, specimens were examined only by outside experts; these instances are clearly noted in the checklist. The names used in this publication generally follow accepted taxonomy and nomenclature (Esslinger 2009) with occasional deviations due to corrections or authors' personal preferences.

 

RESULTS AND DISCUSSION

Over 600 taxa were verified from herbarium collections. A literature review indicated that 195 of these taxa were not previously reported or mapped in the state of Virginia (even under their nomenclatural synonyms). Notably, this total does not include the 52 state records recently reported by Hodkinson & Case (2008). Three of the records reported here represent species new to North America at the time of initial publication (i.e., June 2009). Arthonia anglica Coppins has previously been treated as Arthonia dryadum R.C. Harris & Ladd ined. (2005), but has been found to be indistinguishable from European material (for a full report of this taxon in North America, see Lendemer et al.; in press). Calvitimela talayana (Haugan & Timdal) Andreev was collected and recorded as part of the Shenandoah National Park Rock Outcrop Management Project, but its existence in North America has been verified and is formally published here; for a complete treatment of this species in North America, see Brodo (2009). Entirely unreported from the continent is the lichenicolous fungus Sphaerellothecium coniodes (Nyl.) Cl. Roux & Diederich, a taxon that is presumably not known from any other collections outside of Europe (Ihlen 1998). A more detailed discussion of this S. coniodes record and the locality at which it was collected is provided by Hodkinson (2010). These records likely reflect the paucity of lichen floristic studies. As more specimens are examined and taxa are added to the list, updates will be posted here [http://www.duke.edu/~bph8/VirginiaLichens/], and a partial dynamic checklist (generated from selected herbarium specimens that have been verified and databased at NY) will be accessible at http://sciweb.nybg.org/Science2/hcol/lena/index.asp.

Drastic differences can be seen in the number of species collected in the Mountains (509) versus each of the other physiographic provinces in the state of Virginia (209 in the Coastal Plain and 165 in the Piedmont). The Mountains of Virginia have previously been shown to represent a hotspot of biodiversity for the world (Stein et al. 2000), and a good deal of undiscovered lichen diversity likely remains in this region. However, collection bias surely accounts for some degree of the discrepancy between provinces. Natural vegetation is much scarcer in the Coastal Plain and Piedmont, where most of the land could be considered urban, suburban, or agricultural. This difference in land use has caused collectors to conduct the vast majority of field work in the Mountains. The results of this study highlight the need for more serious collecting in the Coastal Plain and Piedmont of Virginia.

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Figure 1. Virginia’s physiographic provinces as defined in this publication. Cited specimens are listed according to whether they were collected in the Coastal Plain (C), Piedmont (P), or Mountains (M) of Virginia. [Image adapted from http://www.wm.edu/geology/virginia/.]

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While herbarium specimens, and not literature citations, were considered to be the standard for the inclusion of taxa in the Virginia lichen list, a single instance of a potentially misleading set of specimens must be addressed. The label data for Teloschistes exilis (Michaux) Vainio, specimen #91 in the exsiccata "Reliquiae Tuckermaniae," states that it was collected in Alexandria, "Virginia" by Dr. Hale. The collections that Dr. Josiah Hale sent to Dr. Edward Tuckerman (i.e., the specimens that were eventually included in "Reliquiae Tuckermaniae") were primarily from Alexandria, Louisiana (Tucker 1979), where one would expect to find T. exilis. It seems likely that someone misread "LA" as "VA," and created this error that can now be found in numerous herbaria. The specimen is even used as the basis for the inclusion of the Alexandria, Virginia, area on the species' range map by Brodo et al. (2001; I. Brodo, pers. comm.). Assuming that this is an error, T. exilis is excluded from the list published here.

           

CHECKLIST

The following checklist of lichen-forming, lichenicolous, and allied fungi has been compiled exclusively from voucher specimens examined by the authors or specialists associated with the present study. Cited specimens are listed according to the physiographic provinces in which they were collected (C – Coastal Plain; P – Piedmont; M – Mountains; Fig. 1). Two projects for which specimens are not credited to the individual collectors are the Forest Health Management Project (collections credited to FHM) and the Shenandoah National Park Rock Outcrop Management Project (collections credited to 'SNP;' specimens housed at Shenandoah National Park). Samples of unidentified species are included only when they represent otherwise unknown genera in Virginia. Several specimens examined yielded difficult and potentially problematic determinations; these are included with a question mark (?). Lichenicolous fungi are designated by an asterisk (*), while lichen allies are marked with a dagger (). Taxa that are potentially new to Virginia (i.e., for which no other report for the state has been seen) are indicated by a double cross ().

 

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Abrothallus hypotrachynae Etayo & Diederich* – M: Harris 54076 (on Hypotrachyna showmanii; NY) 

Acarospora badiofusca (Nyl.) Th. Fr. – P: Guccion 1083 (NY)

Acarospora fuscata (Schrader) Arnold – C: Hodkinson 2599 (DUKE); M: Harris 26918 (NY), 36708 (NY), SNP 68, 71, 87, 97, 174

Acarospora glaucocarpa (Ach.) Körber – M: Hubricht B388 (US), B408 (US)

Acarospora obpallens (Nyl. ex Hasse) Zahlbr. – M: Ihrman 7422 (Shen.)

Acarospora umbilicata Bagl. ? – M: Flenniken 1466 (NY)

Agonimia opuntiella (Buschardt & Poelt) Vĕzda – M: Guccion 1053 (NY), 1388 (Guccion, pers. herb.)

Ahtiana aurescens (Tuck.) Thell & Randlane – M: Culberson 11429 (DUKE), 11329 (DUKE), Hale 12924 (DUKE)

Alectoria fallacina Mot. – M: Dey 5627 (DUKE)

Allocetraria oakesiana (Tuck.) Randlane & Thell – M: Reed 67471 (NY), 62279 (NY), Hale 12419 (US)

Amandinea milliaria (Tuck.) P. May & Sheard – C: Hodkinson 1341 (DUKE), 2727 (NY)

Amandinea polyspora (Willey) E. Lay & P. May – C: Hodkinson 335 (DUKE), 353 (NY); P: Guccion 1084 (NY); M: Anderson 08June1936 (NY)

Amandinea punctata (Hoffm.) Coppins & Scheid. – C: Hodkinson 309 (DUKE), 1347 (NY), 3259 (DUKE); M: Harris 26861B (NY), 36684 (NY)

Amandinea submontana Marbach – C: Hodkinson 3782 (DUKE)

Anaptychia palmulata (Michx.) Vainio – C: Hodkinson 2317 (DUKE), Wiseman 197 (DUKE); P: Hodkinson 5022 (DUKE), 5023 (DUKE), Hale 14610 (DUKE); M: Dey 5692 (DUKE), 5711 (DUKE), Harris 36629 (NY), Buck 20702 (NY)

Anisomeridium biforme (Borrer) R. C. Harris M: Buck 28406 (NY)

Anisomeridium distans (Willey) R. C. Harris – M: Buck 28349 (NY)

Anisomeridium polypori (Ellis & Everh.) M. E. Barr – P: Guccion 1085 (NY); M: Harris 36692 (NY), Buck 36977 (NY), Lendemer 3802 (NY), 3796 (NY), Brodo 9382 (DUKE)

Anzia colpodes (Ach.) Stizenb. – C: Luttrell 1692 (US); P: Reed 58836 (NY), 63113 (NY), Hale 14609 (DUKE); M: Reed 66901 (NY), Culberson 11416 (DUKE), 11356 (DUKE)

Arctoparmelia centrifuga (L.) Hale – M: SNP 39   

Arthonia anglica Coppins [= Arthonia dryadum R.C. Harris & Ladd ined.] – M: Perlmutter 1350 (NCU)

Arthonia caesia (Flotow) Körber – C: Hodkinson 367 (DUKE); M: Buck 36979 (NY)

Arthonia lapidicola (Taylor) Branth & Rostrup ? – M: Hodkinson 9341 (NY)

Arthonia quintaria Nyl. – C: Hodkinson 2191 (DUKE), 2456 (NY); M: Hodkinson 9249 (NY)

Arthonia rubella (Fée) Nyl. – C: Hodkinson 947 (DUKE)

Arthothelium spectabile A. Massal. – M: Harris 36694 (NY), Hodkinson 9202 (in a mixed collection labeled as 'Pertusaria velata UV+;' NY)

Arthothelium taediosum auct. Amer. – C: Hodkinson 4171 (DUKE); M: Hodkinson 9007 (Hodkinson, pers. herb.), Small 25July1892 (NY)

Aspicilia caesiocinerea (Nyl. ex Malbr.) Arnold – P: Harris 36659 (NY); M: Anderson _June1936 (NY)

Aspicilia cinerea (L.) Körber – P: Guccion 1086 (NY); M: Harris 12930 (NY), SNP 2, 5, 34, 65, 68, 70, 71, 75, 94, 145, 169

Aspicilia laevata (Ach.) Arnold – M: Buck 36966 (NY)

Aspicilia verrucigera Hue – M: Anderson 29June1935 (NY)

Bacidia coprodes (Körber) Lettau – C: Hodkinson 3274 (NY), 4020 (DUKE); M: Hodkinson 9253 (NY)

Bacidia helicospora S. Ekman – C: Hodkinson 3771 (DUKE)

Bacidia heterochroa (Müll. Arg.) Zahlbr. – C: Hodkinson 324 (DUKE), Reed 63970 (NY); M: Harris 36630 (NY)

Bacidia rubella (Hoffm.) A. Massal. – M: Harris 9941 (MIN)

Bacidia schweinitzii (Fr. ex E. Michener) A. Schneider – C: Hodkinson 1867 (NY), 2415 (DUKE), Reed 60641 (NY); P: Reed 127742 (NY), Culberson 6735 (DUKE); M: Lendemer 3829 (NY), Culberson 6735 (DUKE), 11199 (DUKE)

Bacidia suffusa (Fr.) A. Schneider – C: Hodkinson 1463 (DUKE); P: Reed 74137 (NY); M: Perlmutter 1354 (NCU)

Bacidina delicata (Leight.) V. Wirth & Vězda ? – M: Buck 53399 (NY)

Bacidina egenula (Nyl.) Vězda – C: Hodkinson 1998 (DUKE); M: Hodkinson 9117 (NY)

Baeomyces rufus (Hudson) Rebent. – M: Buck 53348 (in a mixed collection labeled as Sphaerellothecium coniodes (Nyl.) Cl. Roux & Diederich; NY)

Bagliettoa baldensis (A. Massal.) Vězda – M: van Ingen _March1920 (NY)

Bathelium carolinianum (Tuck.) R. C. Harris – C: Hodkinson 423 (DUKE)

Biatora appalachensis Printzen & Tønsberg – M: Harris 54023 (NY)

Biatora longispora (Degel.) Lendemer & Printzen – M: Buck 20659 (NY), 20686 (NY)

Biatora printzenii Tønsberg – M: Harris 12892 (NY)

Biatora vernalis (L.) Fr. – M: Buck 53372 (NY)

Bilimbia sabuletorum (Schreber) Arnold – M: Buck 28413 (NY), Harris 12945 (NY), 12962 (NY), Hodkinson 9275 (DUKE)

Botryolepraria lesdainii (Hue) Canals et al. – M : Perlmutter 1368 (NCU)

Bryoria furcellata (Fr.) Brodo & D. Hawksw. – M: Buck 28353 (NY), Culberson 11198 (DUKE), 20123 (DUKE), Hodkinson 8957 (NY), Reed 68927 (NY)

Buellia curtisii (Tuck.) Imshaug – C: Hodkinson 303 (DUKE), 388 (DUKE), 1352 (NY), Reed 118118 (NY), Anderson _June1936 (NY); P: Reed 59121 (NY); M: Harris 36628 (NY)

Buellia dialyta (Nyl.) Tuck. – M: Harris 36779 (NY)

Buellia elizae (Tuck.) Tuck. – C: Tuckerman _1854 (Holotype/Lectotype?; F)

Buellia spuria (Schaerer) Anzi – M: Luttrell 2896 (US)

Buellia stellulata (Taylor) Mudd – M: SNP 60, 75, 266

Buellia stillingiana J. Steiner – C: Hodkinson 947 (DUKE), Reed 46158 (NY); P: Reed 59792 (NY); M: Harris 26894 (NY), Hodkinson 9196 (DUKE)

Buellia vernicoma (Tuck.) Tuck. – M: Anderson 09June1936 (NY)

Buellia wheeleri R.C. Harris (for more information on this taxon's distribution, see Lendemer & Hodkinson 2009a) – C: Hodkinson 10239 (NY)

Bulbothrix confoederata (Culb.) Hale – P: Reed 59656 (NY)

Calicium abietinum Pers. – M: Guccion 1054 (NY)

Calicium glaucellum Ach. – M: Harris 10659 (NY), Buck 20685 (NY)

Caloplaca camptidia (Tuck.) Zahlbr. – P: Reed 62735 (NY)

Caloplaca cerina (Hedwig) Th. Fr. – M: Anderson _June1936 (NY)

Caloplaca chrysophthalma Degel. – M: Harris 36764 (NY), Guccion 1055 (NY)

Caloplaca citrina (Hoffm.) Th. Fr. – M: Harris 12866 (NY)

Caloplaca crenulatella (Nyl.) H. OlivierM: Harris 10678 (NY), Hodkinson 9142 (NY)

Caloplaca flavocitrina (Nyl.) H. Olivier – C: Hodkinson 1026 (originally reported as "Caloplaca citrina (Hoffm.) Th. Fr."  by Hodkinson & Case, 2008; NY)

Caloplaca flavovirescens (Wulfen) Dalla Torre & Sarnth. – C: Hodkinson 2331 (DUKE); M: Buck 2063 (NY), Harris 26967 (NY)

Caloplaca microphyllina (Tuck.) Hasse – M: Anderson _June1936 (NY), Culberson 11285 (DUKE)

Caloplaca oxfordensis Fink – M: Ihrman 1463 (Shen.)

Caloplaca subsoluta (Nyl.) Zahlbr. – C: Hodkinson 1023 (DUKE), 1024 (DUKE), 1025 (NY); P: Guccion 1056 (NY), 1057 (NY); M: Ihrman 1582 (Shen.)

Calvitimela talayana (Haugan & Timdal) Andreev – M: SNP 343

Candelaria concolor (Dickson) Stein – C: Hodkinson 304 (NY), 1918 (DUKE); P: Guccion 1087 (NY), Hodkinson 10266 (fertile; Hodkinson, pers. herb.); M: Hale 14605 (DUKE), Culberson 10109 (DUKE), 11295 (DUKE)

Candelaria fibrosa (Fr.) Müll. Arg. – P: Williams 326 (DUKE); M: Culberson 11166 (DUKE)

Candelariella efflorescens R.C. Harris & Buck – Guccion 1349 (fertile; det. Guccion, not seen; Guccion, pers. herb.).

Candelariella vitellina (Hoffm.) Müll. Arg. – P: Guccion 1058 (NY); M: Lehnert s.n. (US)

Candelariella xanthostigma (Ach.) Lettau – M: Guccion 1059 (NY)

Candelariella xanthostigmoides (Müll. Arg.) R.W. Rogers (previously reported here as 'Candelariella reflexa (Nyl.) Lettau') – C: Hodkinson 88 (NY), 307 (DUKE), 805 (DUKE); P: Hodkinson 10252 (Hodkinson, pers. herb.); M: Harris 54026 (NY)

Canoparmelia caroliniana (Nyl.) Elix & Hale – C: Hodkinson 830 (DUKE), Reed 61403 (NY); P: Reed 59212 (NY); M: Seymour 25July1891 (NY), Hale 83 (DUKE)

Canoparmelia crozalsiana (de Lesd. ex Harm.) Elix & Hale – C: Hodkinson 289 (DUKE), 525, FHM 3707732-7 (DUKE); P: Hodkinson 5020 (DUKE)

Canoparmelia texana (Tuck.) Elix & Hale – C: Hodkinson 505 (NY), 2639 (DUKE); P: FHM 3607862-1 (DUKE)

Catillaria nigroclavata (Nyl.) Schuler – M: Harris 26877A (NY)

Cercidospora sp.* – M: SNP 176 (on Rhizoplaca subdiscrepans)

Cetradonia linearis (Evans) J.-C. Wei & Ahti – M: Wieboldt 9385 (VPI)

Cetraria arenaria Kärnef. – M: Luttrell 3245 (DUKE), Allard 3908 (US), 4665a (US)

Cetrelia chicitae (Culb.) W. L. Culb. & C. Culb. – M: Buck 28338 (NY); Culberson 11423 (DUKE), Hale 78 (DUKE), Harris 36727 (NY), Lendemer 4473 (NY)

Cetrelia monachorum (Zahlbr.) W. L. Culb & C. Culb. – M: Dey 5645 (DUKE), 5665 (DUKE), 5691 (DUKE), 5716 (DUKE), 5725 (DUKE)

Cetrelia olivetorum (Nyl.) W. L. Culb. & C. Culb. – M: Culberson 11180 (DUKE), 11238 (DUKE), 17375 (DUKE), Harris 36755 (NY), Reed 129134 (NY)

Chaenotheca ferruginea (Turner ex Sm.) Mig. – M: Buck 28355 (NY), 20708 (NY)

Chaenotheca furfuracea (L.) Tibell – M: Hodkinson 9050 (DUKE)

Chaenotheca trichialis (Ach.) Th. Fr. – M: Buck 28340 (NY), 28405 (NY)

Chaenothecopsis debilis (Turner & Borrer ex Sm.) Tibell†‡ – M: Lendemer 3825 (PH)

Chaenothecopsis pusilla (Ach.) A. F. W. Schmidt†‡ – M: Buck 2089 (NY)

Chrysothrix onokoensis (Wolle) R.C. Harris & Ladd – M: Slaughter _1975-76 (EIU)

Chrysothrix xanthina (Vain.) Kalb C: Hodkinson 3257 (DUKE); P: Hodkinson 10247 (Hodkinson, pers. herb.); M: Guccion 1060 (NY)

Cladonia apodocarpa Robbins – C: Hodkinson 1946 (DUKE), Torrey _Feb1937 (NY); P: Guccion 1088 (NY), Luttrell 3056 (US); M: Harris 12862 (NY), Buck 36959 (NY)

Cladonia arbuscula (Wallr.) Flotow – M: Culberson 11340 (DUKE)

Cladonia atlantica A. Evans – C: Reed 128998 (NY), Torrey _Jan1937 (NY), _1937 (NY); P: Gray 925 (US), 959 (US)

Cladonia beaumontii (Tuck.) Vainio ? – C: Hodkinson 527 (DUKE)

Cladonia caespiticia (Pers.) Flörke – C: Hodkinson 1628 (NY), 2234 (DUKE); P: Guccion 1089 (NY), Killip 6534 (US); M: Harris 10660 (NY), 26935 (NY), Culberson 11219 (DUKE)

Cladonia cariosa (Ach.) Sprengel – P: Reed 59109 (NY)

Cladonia caroliniana Tuck. – P: Reed 103238 (lacking squamatic acid; NY), Luttrell 2231 (US); M: Luttrell 2423 (US)

Cladonia chlorophaea (Flörke ex Sommerf.) Sprengel – M: Harris 36736 (NY)

Cladonia coccifera (L.) Willd. – M: SNP 30, 115

Cladonia coniocraea (Flörke) Sprengel – C: Torrey _1937 (NY); M: Murrill 491 (NY)

Cladonia crispata (Ach.) Flotow – M: Anderson _1937 (thamnolic acid chemotype; NY), Slaughter _1975-76 (squamatic acid chemotype; EIU)

Cladonia cristatella Tuck. – C: Hodkinson 87 (NY), 639 (DUKE), 1839 (DUKE); P: Allard 10514 (NY), Hale 15112 (DUKE), Hodkinson 5005 (DUKE), 5033 (DUKE); M: Buck 36967 (NY), Dey 5578 (DUKE), Culberson 11379 (DUKE)

Cladonia cryptochlorophaea Asahina – P: Hodkinson 5018 (DUKE), Reed 60367 (NY); M: SNP 122, Leeming 04June1892 (NY)

Cladonia cylindrica (A. Evans) A. Evans – C: Reed 139827 (NY); P: Hodkinson 10240 (NY), 11201 (DUKE); M: Harris 36777 (NY)

Cladonia didyma (Fée) Vainio var. didyma – C: Torrey _Feb1937 (NY); M: Seymour 01Aug1891 (NY)

Cladonia didyma var. vulcanica (Zoll. & Moritzi) Vainio – C: Hodkinson 1005 (DUKE), 2187; M: Reed 111567 (NY), Harris 27236 (NY), 36721 (NY)

Cladonia digitata (L.) Hoffm. – M: Dey 5555 (DUKE), 5652 (DUKE)

Cladonia dimorphoclada Robbins – C: Torrey _1937 (NY); M: Tucker 17796 (NY)

Cladonia farinacea (Vainio) A. Evans – C: Luttrell 3585 (US), Torrey _1937 (US)

Cladonia fimbriata (L.) Fr. – M: Anderson _June1936 (NY), Culberson 19160 (DUKE)

Cladonia floerkeana (Fr.) Flörke – M: Dey 5607A (DUKE)

Cladonia floridana Vainio – C: Luttrell 2054 (US)

Cladonia furcata (Hudson) Schrader – M: Buck 36951 (NY), Harris 36715 (NY), Culberson 20581 (DUKE)

Cladonia grayi G. Merr. ex Sandst. – C: Hodkinson 534 (NY), 653 (DUKE), 688 (DUKE), Reed 60703 (no fumarprotocetraric acid; NY), 62595 (no fumarprotocetraric acid; NY), 66144 (NY); P: Reed 63136 (no fumarprotocetraric acid; NY), 68306 (fumarprotocetraric acid chemotype; NY), 84200 (fumarprotocetraric acid chemotype; NY); M: Harris 26927 (NY), 26885 (NY), SNP 186, Culberson 19642 (DUKE)

Cladonia incrassata Flörke – M: Harris 26964 (NY), 36685 (NY), Dey 5681 (DUKE)

Cladonia innominata Lendemer – P: Reed 138220 (NY); M: Culberson 19161 (DUKE), 19162 (DUKE)

Cladonia leporina Fr. – C: Irwin & Payne 37252A (NY), Walton 17April2002 (NY)

Cladonia macilenta Hoffm. var. macilenta – P: Guccion 1090 (NY); M: SNP 254

Cladonia macilenta var. bacillaris (Genth) Schaerer – C: Hodkinson 610 (DUKE), 1300 (NY), 1324 (DUKE), Anderson _June1936 (NY), Torrey _1937 (NY); P: Guccion 1091 (NY), Luttrell 1674 (US); M: Harris 26851 (NY), 36670 (NY)

Cladonia mateocyatha Robbins – C: Hodkinson 529 (DUKE); P: Luttrell 2996 (US), Reed 59731 (NY); M: Guccion 1092 (NY)

Cladonia merochlorophaea Asahina – M: Dey 5638 (DUKE)

Cladonia ochrochlora Flörke – C: Hodkinson 2245 (DUKE), 2258 (NY), Reed 59046 (NY); P: Hodkinson 5009 (DUKE), Reed 63129 (NY); M: Reed 83277 (NY)

Cladonia parasitica (Hoffm.) Hoffm. – C: Hodkinson 1290 (DUKE); P: Guccion 1093 (NY); M: Culberson 11489 (DUKE), Dey 5635 (DUKE)

Cladonia perlomera Kristinsson – P: Reed 59630 (NY)

Cladonia petrophila R. C. Harris – M: Harris 12928 (NY), SNP 73

Cladonia peziziformis (With.) J. R. Laundon – C: Hodkinson 425 (NY), 1283 (DUKE), Teller & Cummings 22Dec1892 (DUKE); P: Hodkinson 5006 (DUKE), Guccion 1094 (NY); M: Buck 36961 (NY)

Cladonia piedmontensis G. Merr. – C: Hodkinson 644 (NY), 679 (DUKE); P: Gray _Feb1928 (NY); M: Allard 6196 (NY), 4275 (NY)

Cladonia pleurota (Flörke) Schaerer – P: Murrill 500 (NY); M: Harris 26883 (NY)

Cladonia polycarpia G. Merr. [chemotypes of Cladonia subcariosa are maintained as separate species in this list] – C: Torrey _Jan1937 (NY)

Cladonia polycarpoides Nyl. [chemotypes of Cladonia subcariosa are maintained as separate species in this list] – C: Hodkinson 646 (DUKE), 649 (NY), 1314 (DUKE), 2291 (DUKE), Reed 111552 (NY); P: Hodkinson 5016 (DUKE), Reed 63075 (NY); M: Reed 106169 (NY)

Cladonia pyxidata (L.) Hoffm. – P: Killip 6539b (US); M: Guccion 1095 (NY)

Cladonia ramulosa (With.) J. R. Laundon – C: Hodkinson 484 (NY), 2080 (DUKE); M: SNP 231

Cladonia rangiferina (L.) F. H. Wigg. – P: Allard 5914 (NY); M: Allard 4663 (NY)

Cladonia rappii A. Evans – C: Torrey _Jan1937 (NY), _Feb1937 (NY); M: Anderson _1937 (NY)

Cladonia ravenelii Tuck. – C: Hodkinson 566 (NY), 1536 (DUKE), Reed 79295 (NY); P: Hodkinson 6135 (DUKE), Dirig 3060 (NY)

Cladonia rei Schaerer – C: Hodkinson 492 (DUKE), 1739 (NY), 2077 (DUKE; Hodkinson specimens originally reported as "Cladonia subulata (L.) F. H. Wigg." by Hodkinson & Case, 2008); M: Harris 26851A (NY), Culberson 19643 (DUKE)

Cladonia robbinsii A. Evans – P: Cooke 16951 (DUKE), Reed 78222 (NY); M: Reed 75675 (NY)

Cladonia scabriuscula (Delise) Nyl. – C: Torrey _Jan1937 (NY)

Cladonia sobolescens Nyl. ex Vainio [chemotypes of Cladonia subcariosa are maintained as separate species in this list] – C: Hodkinson 561 (DUKE), Torrey _Jan1937 (NY), _Feb1937 (NY); P: Torrey _Feb1937 (NY); M: Allard 6429 (NY)

Cladonia squamosa Hoffm. – C: Hodkinson 728 (NY), 1322 (DUKE), 1489 (DUKE); P: Guccion 1096 (NY); M: Harris 36705 (NY), 36773 (NY), SNP 222

Cladonia strepsilis (Ach.) Grognot – C: Hodkinson 1323 (DUKE), Torrey _1937 (NY), _Feb1937 (NY); P: Torrey _1937 (NY); M: Anderson _June1936 (NY)

Cladonia stygia (Fr.) Ruoss – M: Ahti 42111 (H; det. Ahti, not seen), 42212 (H, US; det. Ahti, not seen)

Cladonia subradiata (Vainio) Sandst. – C: Hodkinson 1775 (DUKE); P: Miller 315 (NY)

Cladonia subtenuis (Abbayes) Mattick – C: Hodkinson 635 (NY), 1984 (DUKE), 2137 (DUKE), Worthley & Guerke 08June1968 (NY); P: Hodkinson 5014 (DUKE), Reed 63131 (NY); M: Allard 4475 (NY), SNP 104, 227

Cladonia uncialis (L.) F. H. Wigg. – M: Allard 10655 (NY), 6033 (NY), 18213 (NY), SNP 55, 229

Cladonia verticillata (Hoffm.) Schaerer – C: Torrey _Feb1937 (NY); P: Allard 6011 (NY); M: Allard 3896 (NY), Culberson 11378 (DUKE)

Coccocarpia erythroxyli (Sprengel) Swinscow & Krog – M: Anderson 09June1936 (NY)

Coccocarpia palmicola (Sprengel) Arv. & D. J. Galloway – P: Reed 129014 (NY); M: Anderson _1937 (NY)

Coenogonium luteum (Dicks.) Kalb & Lücking – M: Hodkinson 9045 (NY)

Coenogonium pineti (Ach.) Lücking & Lumbsch – C: Teller & Cummings 19Dec1892 (NY); M: Buck 39138 (NY), 28346 (NY)

Collema bachmanianum (Fink) Degel. – C: Hodkinson 1364 (DUKE), 2098 (DUKE), 2355 (NY)

Collema coccophorum Tuck. – M: Guccion 1061 (NY)

Collema conglomeratum Hoffm. – P: Luttrell 1602 (US); M: Culberson 11328 (DUKE), 11413 (DUKE)

Collema flaccidum (Ach.) Ach. – P: Miller 515 (US); M: Hermann 14982 (US)

Collema furfuraceum (Arnold) Du Rietz – M: Hermann 14982 (NY), Hale 18190 (US)

Collema fuscovirens (With.) J. R. Laundon M: Culberson 11312 (DUKE)

Collema subflaccidum Degel. – C: Wiseman 190 (DUKE), M: Lendemer 3810 (NY), Harris 36735 (NY), Culberson 11424 (DUKE)

Collema tenax (Sw.) Ach. – M: Hodkinson 9214 (NY)

Conotrema urceolatum (Ach.) Tuck. – M: Harris 26866 (NY), 26989 (NY), Culberson 11224 (DUKE)

Cornutispora lichenicola D. Hawksw. & B. Sutton* – M: Harris 54077 (on Lecanora strobilina; NY)

Cyphelium tigillare (Ach.) Ach. – M: Slaughter _1975-76 (EIU)

Cystobasidium hypogymniicola Diederich & Ahti* – M: Harris 54128 (on Hypogymnia incurvoides; NY), 54129 (on Hypogymnia krogiae; NY)

Dactylospora athallina (Müll. Arg.) Hafellner* – M: Buck 53348 (on Baeomyces rufus; NY) 

Dactylospora glaucomarioides (Willey ex Tuck.) Hafellner* – M: Hodkinson s.n. (on Ochrolechia yasudae; Hodkinson, pers. herb.)

Dactylospora lobariella (Nyl.) Hafellner* – M: Hafellner 2817 (US; det. Hafellner, not seen)

Dermatocarpon arenosaxi Amtoft – M: Wieboldt 7213 (VPI)

Dermatocarpon luridum (With.) J. R. Laundon – P: Guccion 1127 (NY); M: Culberson 11334 (DUKE), Hermann 15145 (DUKE)

Dermatocarpon muhlenbergii (Ach.) Müll. Arg. – M: Culberson 11234 (DUKE), 11309 (DUKE), 11341 (DUKE)

Dibaeis baeomyces (L. f.) Rambold & Hertel – C: Hodkinson 1770 (DUKE), Reed 138768 (NY), 139463 (NY); P: Hodkinson 6083 (DUKE); M: Buck 36973 (NY)

Dictyocatenulata alba Finley & E. F. Morris – M: Hodkinson 8991 (NY)

Dimelaena oreina (Ach.) Norman – P: Guccion 1097 (gyrophoric acid chemotype; NY); M: Harris 26901 (gyrophoric acid chemotype; NY), 13004 (gyrophoric acid chemotype; NY), SNP 34 (gyrophoric acid chemotype), 65 (fumarprotocetraric acid chemotype), 67 (fumarprotocetraric acid chemotype), 155 (fumarprotocetraric acid chemotype), 174 (gyrophoric acid chemotype), 176 (gyrophoric acid chemotype), Culberson 11461 (gyrophoric acid chemotype; DUKE)

Diplolaeviopsis ranula Giralt & D. Hawksw.* – M: Harris 54078 (on Lecanora strobilina; NY), Hodkinson 9024 (on Lecanora strobilina; DUKE) 

Diploschistes muscorum (Scop.) R. Sant. – M: Harris 26858 (NY), Leeming 13June1892 (NY)

Diploschistes scruposus (Schreber) Norman – M: Guccion 106 (NY), SNP 65, 271

Endocarpon diffractellum (Nyl.) Gueidan & Cl. Roux – M: Harris 12935A (NY), 10676 (NY), Hodkinson 9316 (DUKE)

Endocarpon pallidulum (Nyl.) Nyl. – M: Hodkinson 9118 (in a mixed collection filed under Theldium zwackhii; NY), 9273 (NY)

Endocarpon petrolepideum (Nyl.) Hue – M: Buck 28315 (NY), Harris 10686 (NY), Hodkinson 9322 (DUKE), Reed 63008 (NY)

Endococcus perpusillus Nyl.* – M: SNP 100 (on Rhizocarpon geographicum), 128 (on Porpidia tuberculosa)

Endococcus propinquus (Körber) D. Hawksworth* – M: SNP 80 (on Porpidia tuberculosa), 273 (on Porpidia tuberculosa)

Eopyrenula intermedia Coppins – M: Harris 12916 (NY), 12897 (NY), 26995 (NY), Hodkinson 9265 (DUKE)

Ephebe lanata (L.) Vainio – M: Culberson 11468 (DUKE)

Evernia mesomorpha Nyl. – M: Anderson 08June1936 (NY), _June1936 (NY)

Everniastrum catawbiense (Degel.) Hale ex Sipman – M: Harris 26878 (NY), Dey 5594 (DUKE), 5600 (DUKE), 5654 (DUKE)

Fellhanera granulosa R. C. Harris & Lendemer – M: Buck 28322 (NY)

Flavoparmelia baltimorensis (Gyeln. & Fóriss) Hale – C: Hodkinson 1956 (NY), 2401 (DUKE); P: Reed 78230 (NY); M: Harris 12894 (NY), SNP 2

Flavoparmelia caperata (L.) Hale – C: Hodkinson 721 (NY), 2287 (DUKE), Reed 61193 (NY); P: Reed 138223 (NY); M: Reed 67666 (NY), Hale 14661 (US)

Flavopunctelia flaventior (Stirton) Hale – M: Harris 26892 (NY), 26909 (NY), Culberson 11328 (DUKE)

Flavopunctelia soredica (Nyl.) Hale – M: Hale 59630 (US)

Fuscidea appalachensis Fryday – M: SNP 24

Fuscidea arcuatula (Arnold) V. Wirth & Vězda – M: Harris 13001 (NY), SNP 3, 6, 26, 27, 42, 60, 65, 67, 71, 97, 100, 113, 120, 129, 145, 155, 156, 157

Fuscidea pusilla Tønsberg – M: Harris 54032 (NY)

Fuscopannaria leucosticta (Tuck.) P. M. Jørg. – C: Wiseman 154 (DUKE); M: Harris 12954 (NY), Hermann 14839 (US)

Graphis scripta (L.) Ach. – C: Hodkinson 831 (DUKE), 968 (NY), 1018 (DUKE); P: Buck 28310 (NY); M: Harris 26890 (NY), 26958 (NY), Buck 20715 (NY)

Gyalecta jenensis (Batsch) Zahlbr. – M: Green _1884 (NY)

Haematomma persoonii (Fée) A. Massal. – C: Hodkinson 1725 (DUKE), 2521 (DUKE), 2591 (NY), Teller & Cummings 19Dec1892 (NY)

Hertelidea pseudobotryosa R.C. Harris et al. – M: Slaughter _1975-76 (EIU)

Heterodermia albicans (Pers.) Swinscow & Krog – C: Hodkinson 475 (DUKE), 1107 (NY), 1789 (DUKE), 2173 (DUKE)

Heterodermia appalachensis (Kurok.) Culb. – M: Reed 66874 (NY), Culberson 11404 (DUKE)

Heterodermia echinata (Taylor) Culb. – C: Anderson _June1935 (NY); M: Murrill 493 (NY)

Heterodermia erecta Lendemer – M: Hodkinson 9186 (NY)

Heterodermia granulifera (Ach.) Culb. – M: Guccion 1062 (NY), Reed 67649 (NY), 67678 (NY)

Heterodermia hypoleuca (Ach.) Trevisan – C: Anderson 03September1934 (NY); P: Guccion 1098 (NY), Hale 15594 (US), Miller 141 (US), 255 (US); M: Lendemer 3871 (NY), 3857 (NY), FHM 3608073-21 (DUKE), 3707968-4 (DUKE)

Heterodermia leucomela (L.) Poelt – M: Harris 36753 (NY), Culberson 11397 (DUKE)

Heterodermia neglecta Lendemer, R. C. Harris, & Tripp – M: Harris 12984 (norstictic acid chemotype; NY), 13011 (acid deficient chemotype; NY), 36681 (acid deficient chemotype; NY), Culberson 11300 (acid deficient chemotype; DUKE)

Heterodermia obscurata (Nyl.) Trevisan – C: Anderson _June1936 (NY); P: Reed 68490 (NY), Hale 15649 (DUKE); M: Culberson 11172 (DUKE)

Heterodermia speciosa (Wulfen) Trevisan – P: Hodkinson 5024 (DUKE), M: Harris 36669 (NY), 12919 (NY)

Heterodermia squamulosa (Degel.) Culb. – M: Lendemer 3861 (NY), Harris 36672 (NY)

Hyperphyscia adglutinata (Flörke) H. Mayrh. & Poelt – M: Allard 5747a (US)

Hyperphyscia syncolla (Tuck. ex Nyl.) Kalb – C: Hodkinson 2063 (NY), 2352 (DUKE), Reed 128034 (NY)

Hypocenomyce anthracophila (Nyl.) P. James & Gotth. Schneider – M: Harris 10665 (NY)

Hypocenomyce friesii (Ach.) P. James & Gotth. Schneider – M: Harris 36739 (NY)

Hypocenomyce scalaris (Ach.) M. Choisy – P: Guccion 1099 (NY); M: Dirig 3056 (NY), 3057 (NY), 3058 (NY), L-3054 (DUKE)

Hypogymnia incurvoides Rass. – M: Harris 54094 (NY), 54095 (NY), 54096 (NY)

Hypogymnia krogiae Ohlsson – M: Harris 26908 (NY), Dey 5680 (DUKE)

Hypogymnia physodes (L.) Nyl. – M: Harris 26906 (NY), Hodkinson 8995 (DUKE)

Hypogymnia tubulosa (Schaerer) Hav. – M: Buck 20656 (NY)

Hypotrachyna afrorevoluta (Krog & Swinscow) Krog & Swinscow – M: Harris 26854 (NY), 26863 (NY), Guccion 1100 (NY)

Hypotrachyna croceopustulata (Kurok.) Hale – M: Hale 18493 (DUKE), Dey 5581 (DUKE)

Hypotrachyna gondylophora (Hale) Hale – M: Pike 4461 (NY)

Hypotrachyna imbricatula (Zahlbr.) Hale – M: Dey 5677 (DUKE), 5723 (DUKE)

Hypotrachyna livida (Taylor) Hale – C: Hodkinson 886 (DUKE), 901 (NY), 908 (DUKE), Reed 64637 (NY); P: Reed 58834 (NY); M: Reed 66953 (NY), Culberson 11333 (DUKE)

Hypotrachyna lividescens (Kurok.) Hale – M: Harris 54036 (previously reported here as Hypotrachyna producta Hale; NY)

Hypotrachyna oostingii (J. P. Dey) Hale – M: Dey 5613 (DUKE), 5663 (DUKE)

Hypotrachyna osseoalba (Vainio) Park & Hale – C: Anderson _June1935 (NY), Reed 66597 (NY); M: Guccion 1392 (Guccion, pers. herb.)

Hypotrachyna pustulifera (Hale) Skorepa – P: Hale 15680 (US), Reed 138731 (NY), 138274 (NY)

Hypotrachyna revoluta (Flörke) Hale – P: Hale 38424 (NY); M: SNP 29

Hypotrachyna rockii (Zahlbr.) Hale – M: Culberson 11327 (DUKE), Hodkinson 9205 (NY)

Hypotrachyna showmanii Hale – C: Hodkinson 478 (DUKE); M: Harris 54038 (NY), 54039 (NY), 54076 (in a mixed collection labelled as Abrothallus hypotrachynae Etayo & Diederich; NY)

Hypotrachyna taylorensis (M. E. Mitch.) Hale – M: Hale 33415 (NY)

Hypotrachyna thysanota (Kurok.) Hale – M: Dey 5585 (DUKE), 5587 (DUKE)

Hypotrachyna virginica (Hale) Hale – M: Harris 26917 (NY), Dey 5679 (DUKE)

Illosporium carneum Fr. – M: Hodkinson 4732 (on Peltigera praetextata; NY)

Imshaugia aleurites (Ach.) S. F. Meyer – C: Reed 64169 (NY); P: Reed 62378 (NY); M: Reed 67411 (NY), Culberson 20122 (DUKE)

Imshaugia placorodia (Ach.) S. F. Meyer – P: Reed 65241 (NY), 63253 (NY); M: Reed 56499 (NY), Anderson 1937 (NY)

Ionaspis alba Lutzoni – M: Slaughter _1975-76 (EIU)

Ionaspis lacustris (With.) Lutzoni – M: Harris 12926 (NY), Culberson 11472 (DUKE)

Lasallia papulosa (Ach.) Llano – M: Harris 12937 (NY), Hermann 14148 (NY), Imshaug 38328 (NY), SNP 19

Lasallia pensylvanica (Hoffm.) Llano – M: Hermann 14157 (NY), SNP 5, 67, 266

Lecania croatica (Zahlbr.) Kotlov – M: Hodkinson 8973 (NY)

Lecania cuprea (A. Massal.) v. d. Boom & Coppins – C: Hodkinson 562 (DUKE)

Lecanora albella (Pers.) Ach. var. albella – M: Guccion 1063 (NY)

Lecanora albella var. rubescens (Imshaug & Brodo) Lumbsch – M: Guccion 1064 (NY)

Lecanora allophana Nyl. – M: Hale 33368 (US)

Lecanora argentata (Ach.) Malme – C: Hodkinson 824 (NY), 827 (DUKE); M: Small 20July1892 (NY)

Lecanora caesiorubella Ach. subsp. caesiorubella – M: Harris 26928 (NY)

Lecanora cenisia Ach. – M: Harris 36743 (NY)

Lecanora chlarotera Nyl. – C: Hodkinson 312 (DUKE), 414 (DUKE), 3255 (NY), 3258 (DUKE), 3265 (DUKE)

Lecanora cinereofusca H. Magn. – M: Buck 28319 (NY), Harris 26889 (NY), 26947 (NY), Hodkinson 9199 (DUKE)

Lecanora dispersa (Pers.) Sommerf. – C: Hodkinson 1242 (DUKE); P: Guccion 1065 (NY); M: Ihrman 358-R (in a mixed collection labeled as Rhizocarpon rubescens Th. Fr.; Shen.)

Lecanora hybocarpa (Tuck.) Brodo – C: Hodkinson 361 (DUKE), 390 (DUKE), 392 (NY); P: Guccion 1101 (NY); M: Harris 36781 (NY), 36742 (NY), Culberson 11270 (DUKE)

Lecanora imshaugii Brodo – M: Harris 26939 (NY), Buck 20698 (NY)

Lecanora louisianae de Lesd. – C: Hodkinson 362 (NY), 412 (DUKE), 625 (DUKE)

Lecanora muralis (Schaerer) Rabenh. – P: Guccion 1102 (NY)

Lecanora oreinoides (Körber) Hertel & Rambold – M: Ihrman 273-R (Shen.)

Lecanora polytropa (Hoffm.) Rabenh. – M: Harris 26925 (NY)

Lecanora pulicaris (Pers.) Ach. – M: Harris 36752 (NY), 26920 (NY)

Lecanora rugosella Zahlbr. – M: Harris 54107 (NY), 54108 (NY)

Lecanora rupicola (L.) Zahlbr. – M: Anderson _June1936 (NY)

Lecanora strobilina (Sprengel) Kieffer – C: Hodkinson 807 (DUKE), 836 (DUKE), 906 (NY), Reed 121097 (NY); P: Maxon 498 (NY), Luttrell 1577 (US), 1870 (US), Culberson 4712 (DUKE); M: Harris 26891 (NY), SNP 6, Culberson 11232 (DUKE), 11382 (DUKE)

Lecanora subimmergens Vainio – M: Harris 12884 (NY)

Lecanora subpallens Zahlbr. – C: Hodkinson 910 (DUKE), 913 (DUKE), 1350 (NY), 1616 (DUKE), Reed 117289 (NY), 118117 (NY); M: Harris 36701 (NY)

Lecanora symmicta (Ach.) Ach. – M: Harris 54109 (NY)

Lecanora thysanophora R. C. Harris – M: Harris 26957 (NY), 36670A (NY)

Lecanora valesiaca (Müll. Arg.) Stizenb. – M: Anderson _1937 (NY), Guccion 1066 (NY), Reed 75677 (NY)

Lecanora wisconsinensis H. Magn. – M: Harris 26881 (NY), Buck 20650 (NY), 20679 (NY)

Lecidea ahlesii (Körber) Nyl. var. ahlesii – M: Perlmutter 1283 (NCU)

Lecidea atrobrunnea (Lam. & DC.) Schaerer [substance-deficient chemotype] – M: Anderson _June1936 (NY)

Lecidea berengeriana (A. Massal.) Th. Fr. – M: Lendemer 3872 (NY), Harris 36675 (NY), Buck 28394 (NY)

Lecidea cyrtidia Tuck. – P: Guccion 1105 (NY); M: Buck 20669 (NY), 20712 (NY), Hodkinson 9341 (in a mixed collection labeled as Arthonia lapidicola ?; NY)

Lecidea deminutula H. Magn. – M: Brodo 9465 (CANL)

Lecidea lapicida (Ach.) Ach. – M: Anderson 08June1935 (NY)

Lecidea plebeja Nyl. – C: Hodkinson 383 (DUKE); M: Harris 10658 (NY)

Lecidea tessellata Flörke – M: SNP 179

Lecidella enteroleucella (Nyl.) Hertel – M: Slaughter _1975-76 (EIU)

Leimonis erratica (Körber) R. C. Harris & Lendemer – P: Luttrell 1572 (US); M: SNP 81

Leiorreuma explicans (Fink) Lendemer – C: Hodkinson 967 (DUKE)

Leiorreuma sericeum (Eschw.) Staiger – C: Hodkinson 565 (DUKE), 978 (NY)

Lempholemma polyanthes (Bernh.) Malme – M: Harris 9956 (NY)

Lepraria adhaerens K. Knudsen, Elix & Lendemer – M: Reed 62274 (NY)

Lepraria caesiella R. C. Harris – C: Hodkinson 2281 (DUKE); P: Reed 60677 (NY); M: Harris 36700 (NY)

Lepraria caesioalba (de Lesd.) J. R. Laundon [fumarprotocetraric acid chemotype] – P: Guccion 1103 (NY); M: SNP 34, 44, 65, 80, 131

Lepraria crassissima (Hue) Lettau – M: SNP 29, 31, 131

Lepraria lobificans Nyl. – C: Hodkinson 1020 (DUKE); M: Lendemer 3799 (NY), 3837 (NY), SNP 270

Lepraria neglecta (Nyl.) Erichsen – M: SNP 1, 4, 73, 87

Lepraria normandinoides Lendemer & R. C. Harris – M: Harris 36656 (NY), 36718 (NY), Hodkinson 8978 (Hodkinson, pers. herb.)

Leptogium austroamericanum (Malme) C. W. Dodge – P: Hale 17924 (US); M: Hale 33249 (US), 12777 (US), 33169 (US)

Leptogium chloromelum (Sw.) Nyl. – M: Anderson 08June1936 (NY), Reed 62190 (NY)

Leptogium corticola (Taylor) Tuck. – C: Wiseman 180 (DUKE); M: Buck 20691 (NY), 28400 (NY)

Leptogium cyanescens (Rabenh.) Körber – C: Hodkinson 424 (NY), 1000 (DUKE); P: Hermann 15002 (NY); M: Buck 20705 (NY), Culberson 11440 (DUKE), Hale 33052 (US), Harris 36730 (NY)

Leptogium dactylinum Tuck. – M: Buck 28409 (NY), Guccion 04Nov1990 (NY), Hodkinson 9185 (Hodkinson, pers. herb.)

Leptogium hirsutum Sierk – M: Guccion 1067 (NY), Murrill 479 (NY)

Leptogium lichenoides (L.) Zahlbr. – M: Harris 26962 (NY), 36728 (NY)

Leptogium marginellum (Sw.) Gray – C: Reed 75743 (NY)

Leptogium millegranum Sierk – P: Hale 14601 (DUKE)

Leptogium tenuissimum (Dickson) Körber* – M: Guccion 158 (NY)

Lichenochora obscuroides (Lindsay) Triebel & Rambold – M: Hodkinson 9349 (on Phaeophyscia sp.; NY)

Lichenoconium erodens M. S. Christ. & D. Hawksw.* – C: Hodkinson 3264 (on Lecanora strobilina; DUKE); M: Buck 28373 (on Punctelia rudecta; NY)

Lichenoconium lecanorae (Jaap) D. Hawksw.* – P: Guccion 1089 (on Cladonia caespiticia; NY)

Lichenostigma cosmopolites Hafellner & Calat.* – M: Slaughter _1975-76 (on Xanthoparmelia conspersa; EIU)

Lichenothelia sp. †‡ M: SNP 2, 6

Lithothelium hyalosporum (Nyl.) Aptroot – M: Wetmore 15407 (MIN)

Lobaria pulmonaria (L.) Hoffm. – M: Reed 67693 (NY), 67622 (NY), Harris 36711 (NY)

Lobaria quercizans Michx. – C: Hodkinson 1013 (DUKE), Reed 63614 (NY), Wiseman 170 (DUKE), 183 (DUKE); P: Reed 60209 (NY), 57537 (NY), 59722 (NY); M: Lendemer 3798 (NY), Harris 36747 (NY)

Lobaria ravenelii (Tuck.) Yoshim. – C: Culberson 16018 (DUKE)

Lobaria scrobiculata (Scop.) DC. – M: Luttrell 2351 (US)

Loxospora ochrophaea (Tuck.) R. C. Harris – M: Leeming 28May1892 (NY), 26June1892 (NY), Hodkinson 9141 (DUKE)

Loxospora pustulata (Brodo & Culb.) R. C. Harris – C: Hodkinson 401 (DUKE), 1010 (DUKE), 1431 (NY); P: Guccion 1104 (NY), Leonard 18737 (US); M: Culberson 11230 (DUKE), 11271 (DUKE)

Marchandiomyces lignicola Lawrey & Diederich – M: Lawrey 1716 (Holotype; NY)

Maronea polyphaea H. Magn. – M: Hodkinson 9285 (NY).

Megalospora porphyritis (Tuck.) R. C. Harris – C: Reed 60718 (NY)

Melanelia culbersonii (Hale) Thell – M: SNP 56, 266

Melanelia stygia (L.) Essl. – M: SNP 84, 88, Anderson 08June1935 (NY)

Melanelixia subaurifera (Nyl.) O. Blanco et al. – M: Reed 66946 (NY), Culberson 11159 (DUKE)

Melanohalea halei (Ahti) O. Blanco et al. – M: Buck 20666 (NY), Culberson 11479 (DUKE)

Melaspilea sp. – M: Perlmutter 1284 (NCU)

Menegazzia subsimilis (H. Magn.) R. Sant. – M: Guccion 1068 (NY), 1069 (NY), Anderson 09June1936 (NY), Culberson 11331 (DUKE), Culberson 11249 (DUKE), 11329 (DUKE), Hale 18236 (US), 33225 (US)

Micarea bauschiana (Körb.) V. Wirth & Vězda – M: Buck 53415 (pale form; NY) 

Micarea lithinella (Nyl.) Hedl. – M: Slaughter _1975-76 (filed as Porpidia crustulata; EIU)

Micarea micrococca (Körber) Gams ex Coppins – M: Hodkinson 9250 (NY)

Micarea peliocarpa (Anzi) Coppins & R. Sant. – M: Buck 28402 (NY), 36972 (NY), Harris 36678 (NY), 36778 (NY), Hodkinson 9000 (Hodkinson, pers. herb.)

Micarea prasina Fr. – M: Harris 12942 (NY), 36680 (NY), 36759 (NY)

Microcalicium arenarium (A. Massal.) Tibell (†/*)‡ – M: SNP 133 (on Psilolechia lucida)

Minutoexcipula mariana V. Atienza* – C: Hodkinson 3251 (on Pertusaria xanthodes; NY), 3252 (on Pertusaria sp.; DUKE)

Minutoexcipula tuckerae V. Atienza & D. Hawksw.* – P: Guccion 1112 (on Pertusaria tetrathalamia s. lat.; NY)

Miriquidica leucophaea (Flörke ex Rabenh.) Hertel & Rambold – M: Anderson 08June1935 (NY)

Muellerella lichenicola (Sommerf. ex Fr.) D. Hawksw.* – M: Hodkinson 9266 (on Caloplaca flavovirescens; NY), 9337 (on Physcia pumilior; NY)

Muellerella pygmaea (Körber) D. Hawksw.* – M: SNP 113 (on Rhizocarpon rubescens)

Mycocalicium albonigrum Nyl. Fink – M: Buck 53306 (NY)

Mycocalicium subtile (Pers.) Szatala – M: Harris 10648 (NY), 36757 (NY), Buck 28395 (NY), 28404 (NY), 36949 (NY)

Mycoporum pycnocarpoides Müll. Arg. – M: Hodkinson 9287 (NY)

Myelochroa aurulenta (Tuck.) Elix & Hale – C: Hodkinson 473 (NY), 2434 (DUKE), Reed 61833 (NY); P: Reed 138275 (NY); M: Lendemer 3816 (NY), Hale 15881 (DUKE)

Myelochroa galbina (Ach.) Elix & Hale – C: Reed 83295 (NY); P: Reed 63086 (NY); M: Lendemer 3849 (NY), Culberson 11247 (DUKE)

Myelochroa obsessa (Ach.) Elix & Hale – P: Reed 68773 (NY); M: Anderson 29June1935 (NY), Culberson 11358 (DUKE)

Nadvornikia sorediata R. C. Harris – C: Hodkinson 563 (DUKE); P: Hodkinson 10245 (Hodkinson, pers. herb.), 10258 (NY); M: Buck 41816 (NY)

Nectriopsis rubefaciens (Ellis & Everh.) M. S. Cole & D. Hawksw.* – M: Harris 26868 (on Aspicilia sp.; NY)

Nephroma bellum (Sprengel) Tuck. – M: Small 16June1892 (NY)

Nephroma helveticum Ach. – C: Luttrell 1766 (US), 2107(US); M: Poff 179 (DUKE), FHM 3707973-7A (DUKE), Hale 12663 (US)

Nephroma parile (Ach.) Ach. – M: Culberson 11453 (NY)

Normandina pulchella (Borrer) Nyl. – M: Buck 20706A (NY), 20705A (NY), 28372 (NY), Harris 12995A (NY)

Ochrolechia africana Vainio – C: Hodkinson 300 (NY), 301 (DUKE), 1945 (DUKE), 2010 (DUKE)

Ochrolechia arborea (Kreyer) Almb. – M: Harris 36763 (NY), Buck 28401 (NY)

Ochrolechia mexicana Vainio – M: Guccion 1070 (NY), Hodkinson 9046 (DUKE)

Ochrolechia pseudopallescens Brodo – C: Reed 63409 (NY), Anderson _June1936 (NY)

Ochrolechia trochophora (Vainio) Oshio – M: Harris 12952 (NY), 26971 (NY), 26988 (NY), 36776 (NY), Buck 28324 (NY), 28356 (NY)

Ochrolechia yasudae Vainio – P: Guccion 1106 (NY); M: Harris 10669 (NY), 36716 (NY), 36725 (NY), SNP 7, 259

Opegrapha gyrocarpa Flotow ? – M: SNP 148

Opegrapha varia Pers. – M: Lendemer 3827 (NY)

Opegrapha vulgata Ach. – C: Hodkinson 397 (DUKE), 2151 (NY); M: Harris 12880 (NY)

Pannaria lurida ssp. quercicola P. M. Jørg. – C: Reed 9099 (Holotype in US; Isotype in NY); P: Hale 14568 (US), Luttrell 4889 (US)

Pannaria subfusca P. M. Jørg.P: Luttrell 3801 (US); M: Culberson 11211 (DUKE), Luttrell 537 (US), 3392 (US), 3557 (US)

Pannaria tavaresii P. M. Jørg. – C: Luttrell 1767 (US), 1922 (US), 2028 (US), 2109 (US)

Parmelia neodiscordans Hale – M: Guccion 1071 (NY)

Parmelia omphalodes (L.) Ach. – M: Anderson 08June1935 (NY), 08June1936 (NY), SNP 110, Hale 88 (DUKE), 14679 (US)

Parmelia saxatilis (L.) Ach. – P: Miller 234 (US); M: Culberson 11181 (DUKE), FHM 3708026-18 (DUKE), Allard 2445 (US), 8324 (US)

Parmelia squarrosa Hale – P: Reed 138258 (NY); M: Reed 79487 (NY), Culberson 11331 (DUKE)

Parmelia sulcata Taylor – P: Reed 63067 (NY); M: Reed 127182 (NY), Culberson 11243 (DUKE)

Parmeliella appalachensis P. M. Jørg – M: Culberson 11495 (DUKE)

Parmelinopsis horrescens (Taylor) Elix & Hale – C: Reed 61032 (NY); M: Harris 36770 (NY), Dey 5573 (DUKE)

Parmelinopsis minarum (Vainio) Elix & Hale – C: Reed 61026 (NY); P: Reed 138728 (NY); M: Lendemer 3831 (NY), Culberson 11430 (DUKE)

Parmelinopsis spumosa (Asahina) Elix & Hale – C: FHM 3707567-17 (DUKE)

Parmeliopsis subambigua Gyelnik – C: Reed 83922 (NY), FHM 3607668-8 (DUKE); P: Reed 138720 (NY), Hale 14595 (DUKE)

Parmotrema arnoldii (Du Rietz) Hale – M: Dey 5629 (DUKE)

Parmotrema austrosinense (Zahlbr.) Hale – C: Hodkinson 1937 (DUKE), 3262 (NY)

Parmotrema cetratum (Ach.) Hale – C: Reed 59393 (NY); P: Reed 63461 (NY); M: Culberson 12892 (DUKE)

Parmotrema crinitum (Ach.) M. Choisy – P: Guccion 1107 (NY); M: Harris 36719 (NY), 26919 (NY), Hale 17783 (DUKE)

Parmotrema despectum Kurok. – C: Reed 61854 (NY), 59407 (NY); P: Reed 63077 (NY)

Parmotrema gardneri (C. W. Dodge) Sérus. – C: Hodkinson 846 (DUKE), 1017 (NY), Reed 138180 (NY); P: Guccion 1108 (NY), Reed 83336 (NY); M: Reed 67674 (NY)

Parmotrema hypoleucinum (Steiner) Hale – C: Hodkinson 621 (originally reported as "Parmotrema hypotropum (Nyl.) Hale" by Hodkinson & Case, 2008; NY), 1857 (DUKE), Reed 64170 (NY), 57542 (NY)

Parmotrema hypotropum (Nyl.) Hale – C: Hodkinson 1310 (DUKE), 1859 (DUKE), Reed 61018 (NY); P: Hodkinson 5010 (DUKE), 5013 (DUKE), Reed 99334 (NY); M: Lendemer 3813 (NY)

Parmotrema louisianae (Hale) Hale – C: Hodkinson 1723 (DUKE), 1723B (NY)

Parmotrema margaritatum (Hue) Hale – P: Reed 63308 (NY); M: Harris 9925 (NY), Hale 18263 (US)

Parmotrema perforatum (Jacq.) A. Massal. – C: Hodkinson 711 (NY), 1910 (DUKE), 2134 (DUKE), Reed 66132 (NY); P: Hodkinson 5027 (DUKE), 5028 (DUKE), Reed 138225 (NY); M: Reed 57756 (NY)

Parmotrema perlatum (Huds.) M. Choisy – M: Reed 84110 (NY), Culberson 11153 (DUKE)

Parmotrema praesorediosum (Nyl.) Hale – C: Hodkinson 1271 (DUKE)

Parmotrema reticulatum (Taylor) M. Choisy – C: Hodkinson 1822 (NY), 2645 (DUKE), Reed 61029A (NY); P: Hodkinson 5011 (DUKE), 5030 (DUKE), Reed 75814 (NY); M: Hodkinson 9201 (apothecia present; Reed 66850 (NY)

Parmotrema stuppeum (Taylor) Hale – M: Harris 36750 (NY), 36761 (NY)

Parmotrema subisidiosum (Müll. Arg.) Hale – C: Hodkinson 290 (NY), 1263 (DUKE), Reed 139462 (NY); P: Hodkinson 5007 (DUKE), Reed 75840 (NY); M: Harris 67668 (NY)

Parmotrema submarginale (Michaux) DePriest & B. Hale – C: Hodkinson 907 (DUKE), 2213 (NY), Reed 128048 (NY); P: Reed 58835 (NY); M: Hale 12891 (DUKE)

Parmotrema subrigidum Egan – C: Reed 63971 (NY)

Parmotrema subtinctorium (Zahlbr.) Hale – P: Reed 138226 (NY); M: Reed 66702 (NY)

Parmotrema tinctorum (Delise ex Nyl.) Hale – C: Luttrell 1709 (US)

Parmotrema ultralucens (Krog) Hale – C: Culberson 16019 (DUKE)

Parmotrema xanthinum (Müll. Arg.) Hale – C: Reed 9097A (NY); M: Seymour 27July-01August1891 (NY)

Peltigera canina (L.) Willd. – M: Allard 3966 (US), 14April1935 (US), Hale 14842 (US)

Peltigera didactyla (With.) J. R. Laundon – M: Luttrell 2426 (US), Anderson 08June1936 (NY), Leeming 04June1892 (NY)

Peltigera elisabethae Gyelnik – M: Culberson 11392 (DUKE)

Peltigera evansiana Gyelnik – M: Guccion 1072 (NY)

Peltigera horizontalis (Hudson) Baumg. – C: Hodkinson 1091 (DUKE); M: Small 5269 (NY), Dey 5704 (DUKE), Hale 18298 (US)

Peltigera hydrothyria Miadlikowska & Lutzoni – M: Allard 4601 (NY), Culberson 11439 (DUKE), 11441 (DUKE)

Peltigera leucophlebia (Nyl.) Gyelnik – M: Murrill 470 (NY), Anderson 09June1936 (NY), Culberson 11313 (DUKE), Allard 4833 (US)

Peltigera neckeri Hepp ex Müll. Arg. M: Hale 10432 (US), 18118 (US), 18181 (US), 18877 (US)

Peltigera neopolydactyla (Gyelnik) Gyelnik – C: Hodkinson 1090 (DUKE), 1745 (NY); M: Hale 17810 (US), 17840 (US)

'Peltigera neopolydactyla sensu lato' – C: Hodkinson 1121 (DUKE), 1122 (DUKE), 1124 (DUKE), 1125 (DUKE), 1128 (NY)

Peltigera phyllidiosa Goffinet & Miadlikowska – C: Hodkinson 2256 (DUKE), Wiseman 177 (DUKE); P: Hodkinson 5025 (DUKE); M: Culberson 11209 (DUKE), Dey 5689 (DUKE), Hodkinson 9200 (NY)

Peltigera polydactylon (Necker) Hoffm. – M: Seymour 27July-01Aug1891 (NY), Culberson 11380 (DUKE), Allard 4086 (US)

Peltigera praetextata (Flörke ex. Sommerf.) Zopf – C: Hodkinson 2168 (DUKE), Wiseman 182 (DUKE); M: Allard 5881 (US), Hodkinson 4732 (NY), Harris 26996 (NY), 36634 (NY), 26942 (NY), Leeming 13June1892 (NY), Anderson 29June1935 (NY), 09June1936 (NY), Culberson 11298 (DUKE), 11305 (DUKE), 11347 (DUKE)

Peltigera rufescens (Weiss) Humb. – C: Hodkinson 1267 (DUKE); M: Allard 2411 (US), 4096 (US), Luttrell 80 (US), 2999 (US)

Peltula sp. – P: Hodkinson 10205 (DUKE, NY)

Pertusaria amara (Ach.) Nyl. – P: Luttrell 2271 (DUKE); M: Buck 28339 (NY), Harris 36783 (NY)

Pertusaria andersoniae Lendemer – M: Anderson 08June1935 (NY)

Pertusaria epixantha R. C. Harris – C: Hodkinson 339 (originally reported as "Pertusaria texana Müll. Arg." by Hodkinson & Case, 2008; DUKE), 389 (originally reported as "Pertusaria texana Müll. Arg." by Hodkinson & Case, 2008; NY), 1201 (DUKE), 3647 (NY); P: Hodkinson 10246 (Hodkinson, pers. herb.); M: Harris 12904 (NY), 12909B (NY), 12911 (NY)

Pertusaria globularis (Ach.) Tuck. – M: Buck 28407 (NY), Hodkinson 9184 (DUKE), Reed 106138 (NY)

Pertusaria hypothamnolica Dibben – P: Reed 62889 (NY)

Pertusaria macounii (Lamb) Dibben – M: Harris 26865 (NY), 26946 (NY), 26952 (NY), 36787 (NY)

Pertusaria multipunctoides Dibben – C: Hodkinson 680 (DUKE), Luttrell 2062 (DUKE); M: Buck 28342 (NY), Schallert 03Apr1938 (NY)

Pertusaria neoscotica Lamb – P: Reed 63455 (NY); M: Harris 26941 (NY), Hodkinson 9182 (NY), Luttrell 2583 (DUKE)

Pertusaria ophthalmiza (Nyl.) Nyl. – C: Anderson 03Sept1934 (NY), Reed 63615 (NY); M: Harris 36771 (NY), 36691 (NY)

Pertusaria ostiolata Dibben – C: Luttrell 1913 (DUKE); P: Guccion 1110 (NY)

Pertusaria paratuberculifera Dibben – C: Hodkinson 971 (DUKE), 1829 (NY), 2165 (DUKE), Reed 117323 (NY), Anderson _June1935 (NY); P: Harris 36660 (NY), Reed 83613 (NY); M: Buck 36968 (NY), Tucker 17738 (NY)

Pertusaria plittiana Erichsen – M: Harris 10662 (NY), 36712 (NY), Guccion 120 (NY), Schallert 1132 (NY)

Pertusaria propinqua Müll. Arg. – C: Hodkinson 1016 (DUKE), Luttrell 1704 (DUKE), 1915 (DUKE)

Pertusaria pustulata (Ach.) Duby – C: Hodkinson 818 (NY), 1402 (DUKE), Luttrell 1947 (DUKE); P: Guccion 1111 (NY); M: Buck 36978 (NY), Guccion 119 (NY), Harris 12875 (NY), 26972 (NY), 27205 (NY)

Pertusaria rubefacta Erichsen – C: Hodkinson 968 (DUKE); M: Buck 28371 (NY), Harris 12970 (NY), 26960 (NY)

Pertusaria shenandoensis Hale & Dibben – M: Dibben 22437 (DUKE), Hale 01June1971 (Holotype; US)

Pertusaria sinusmexicani Dibben – C: Luttrell 1771 (DUKE)

Pertusaria subpertusa Brodo – C: Hodkinson 399 (DUKE), 508 (NY), Luttrell 1604 (DUKE); P: Guccion 1109 (NY)

Pertusaria tetrathalamia (Fée) Nyl. – C: Luttrell 2038 (DUKE)

'Pertusaria tetrathalamia sensu lato' [the thallus xanthone is lichexanthone not coronatone; may be an undescribed species] P: Guccion 1112 (NY)

Pertusaria texana Müll. Arg. – P: Luttrell 1562 (DUKE)

Pertusaria trachythallina Erichsen – C: Luttrell 1638 (DUKE), 1695 (DUKE); M: Harris 36785 (NY), 12977 (NY), 26855 (NY), 26974 (NY), Reed 106142 (NY)

Pertusaria valliculata Dibben – P: Guccion 1113 (NY)

Pertusaria velata (Turner) Nyl. – C: Luttrell 1912 (DUKE), 1916 (DUKE); P: Luttrell 2236 (DUKE); M: Harris 26940 (NY), 36693 (NY), Hodkinson 9181 (UV-; Hodkinson, pers. herb.), 9202 (UV+; NY), Lendemer 3797 (NY)

Pertusaria waghornei Hulting – M: Anderson _June1936 (NY), Luttrell 2657 (DUKE)

Pertusaria xanthodes Müll. Arg. – C: Hodkinson 915 (NY), 2095 (DUKE); P: Guccion 1114 (NY)

Phaeocalicium polyporaeum (Nyl.) Tibell†‡ – M: Harris 36749 (NY), Miller 24254 (NY)

Phaeographis inusta (Ach.) Müll. Arg. – C: Anderson 03Sept1934 (NY), Hodkinson 810 (NY), 933 (DUKE), 1130 (DUKE), 1434 (DUKE); P: Hodkinson 10256 (Hodkinson, pers. herb.)

Phaeophyscia adiastola (Essl.) Essl. – C: Hodkinson 833 (DUKE), Anderson _June1935 (NY); M: Buck 36969 (NY), Guccion 1075 (NY), Harris 36653 (NY), Hodkinson 9021 (fertile; DUKE)

Phaeophyscia ciliata (Hoffm.) Moberg – C: Reed 62010 (NY); M: Culberson 11286 (in a mixed collection labeled as Phaeophyscia hirtella Essl.; DUKE), Guccion 1115 (NY)

Phaeophyscia hirsuta (Mereschk.) Essl. – C: Hodkinson 842 (DUKE), 1530 (NY); P: Guccion 17May1998 (DUKE); M: Guccion 1073 (NY)

Phaeophyscia hirtella Essl. – C: Hodkinson 460 (DUKE), 1612 (NY); M: Culberson 11286 (DUKE)

Phaeophyscia hispidula (Ach.) Essl. – M: Small 04July1892 (NY)

Phaeophyscia pusilloides (Zahlbr.) Essl. – P: Guccion 1116 (NY), Allard 5747 (US), 5749 (US); M: Harris 12943 (NY), Culberson 11337 (DUKE)

Phaeophyscia rubropulchra (Degel.) Essl. – C: Hodkinson 375 (NY), 1515 (DUKE); P: Guccion 1117 (NY), Allard 5747b (US); M: Harris 36698 (NY), 36663 (NY)

Phaeophyscia sciastra (Ach.) Moberg – M: Reed 58343 (NY)

Phaeophyscia squarrosa Kashiw. – P: Guccion 1118 (NY); M: Guccion 1400 (Guccion, pers. herb.)

Phaeospora sp.* – M: Slaughter _1975-76 (on an unknown host originally identified as 'Heterocarpon ochroleucum;' EIU)

Phaeosporobolus alpinus R. Sant., Alstrup & D. Hawksw.* – P: Hodkinson 10257 (on Pertusaria sp.; NY); M: Ihrman 1694 (on Ochrolechia arborea; Shen.)

Phlyctis argena (Sprengel) Flotow – M: Harris 26951 (NY), Buck 20714 (NY), 20722 (NY)

Phlyctis ludoviciensis (Müll. Arg.) Lendemer – P: Reed 138730 (NY)

Phlyctis petraea R. C. Harris ined.M: Harris 36724 (NY), SNP 1, 131

Phyllopsora corallina (Eschw.) Müll. Arg. – C: Hodkinson 1483 (DUKE)

Physcia americana G. Merr. – C: Anderson _June1935 (NY), Reed 68548 (NY); P: Hale 14612 (DUKE); M: Harris 12914 (NY), Lendemer 3860 (NY)

Physcia millegrana Degel. – C: Hodkinson 288 (DUKE), 2100 (NY), Anderson 03Sept1934 (NY); P: Culberson 4671 (DUKE); M: Harris 26978A (NY)

Physcia phaea (Tuck.) J. W. Thomson – M: Harris 12914 (NY), 26937 (NY)

Physcia pseudospeciosa J. W. Thomson – M: Guccion 1074 (NY), Hale 17910 (US), 18151 (US)

Physcia pumilior R. C. Harris – C: Hodkinson 1129 (DUKE), 1343 (NY); P: Guccion 1119 (NY); M: Perlmutter 1296 (NCU)

Physcia sorediosa (Vainio) Lynge – C: Reed 62011 (NY); M: Harris 12913 (NY)

Physcia stellaris (L.) Nyl. – C: Hodkinson 2505 (DUKE); P: Buck 30957 (NY), Guccion 1120 (NY); M: Harris 26922 (NY), Allard 5742d (US)

Physcia subtilis Degel. – C: Hodkinson 1273 (NY), 1280 (DUKE); P: Guccion 1121 (NY); M: Buck 20675 (NY), Harris 36664 (NY), SNP 19

Physciella chloantha (Ach.) Essl. – C: Anderson 21June1988 (NY), Hodkinson 501 (DUKE), 1466 (NY); P: Hodkinson 10253 (Hodkinson, pers. herb.)

Physciella nepalensis (Poelt) Essl. – M: Ihrman 963 (Shen.)

Physconia detersa (Nyl.) Poelt – M: Culberson 11283 (DUKE), Hodkinson 9281 (NY)

Physconia leucoleiptes (Tuck.) Essl. – C: Hodkinson 2001 (lacking gyrophoric acid; DUKE); M: Harris 36662 (lacking gyrophoric acid; NY), 36775 (lacking gyrophoric acid; NY), Hale 33333 (gyrophoric acid chemotype = P. kurokawae; US), 33355 (gyrophoric acid chemotype = P. kurokawae; US)

Placidiopsis minor R. C. Harris – M: Harris 9965 (NY)

Placidium arboreum (Schw. ex E. Michener) Lendemer – C: Wiseman 184 (DUKE); M: Purcell 3853 (NY), Culberson 11410 (DUKE), Luttrell 4180 (US)

Placidium lacinulatum (Ach.) Breuss – P: Reed 68770 (NY)

Placidium squamulosum (Ach.) Breuss – M: Guccion 1076 (NY)

Placynthiella icmalea (Ach.) Coppins & P. James – P: Hodkinson 10254 (DUKE); M: Buck 28328 (NY), Harris 26858A (NY)

Placynthiella oligotropha (J. R. Laundon) Coppins & P. James – M: Buck 20681 (NY)

Placynthium nigrum (Hudson) Gray – M: Harris 10681 (NY), 10679A (NY)

Platismatia tuckermanii (Oakes) W. L. Culb. & C. Culb. – P: Reed 83337 (NY); M: Reed 66817 (NY), Culberson 11251 (DUKE)

Polysporina simplex (Davies) Vězda – C: Luttrell 2081 (US); P: Luttrell 1575 (US); M: Ihrman 1601 (Shen.)

Porina heterospora (Fink) R. C. Harris – C: Hodkinson 2244 (DUKE)

Porpidia albocaerulescens (Wulfen) Hertel & Knoph – P: Guccion 1122 (NY); M: Culberson 11316 (DUKE), Harris 26916 (NY), 12924 (NY), 36633 (NY), SNP 271

Porpidia crustulata (Ach.) Hertel & Knoph – M: Harris 9955 (NY), 36638 (NY), 38695 (NY)

Porpidia lowiana Gowan ? – M: SNP 80, 128

Porpidia macrocarpa f. nigrocruenta (Anzi) Fryday – M: Anderson 09June1936 (NY), Harris 36765 (NY)

Porpidia subsimplex (H. Magn.) Fryday – M: Harris 26992 (NY), Luttrell 2758 (US), SNP 64

Porpidia tuberculosa (Sm.) Hertel & Knoph – M: SNP 80, 81, 97, 113, 128, 129, 145, 270

Protoblastenia rupestris (Scop.) J. Steiner – M: Harris 12863 (NY)

Pseudevernia cladonia (Tuck.) Hale & W. L. Culb. – M: Harris 26912 (NY), Buck 53401 (NY)

Pseudevernia consocians (Vainio) Hale & W. L. Culb. – M: Harris 26857 (NY)

Pseudocyphellaria aurata (Ach.) Vainio – C: Reed 68617 (NY)

Pseudocyphellaria crocata (L.) Vainio – M: Seymour 27July-01Aug1891 (NY), Leeming 26June1892 (NY)

Pseudosagedia cestrensis (Michener) R. C. Harris – C: Hodkinson 997 (DUKE); P: Buck 28313 (NY); M: Harris 12932 (NY)

Pseudosagedia guentheri (Flotow) Hafellner & Kalb – M: Lendemer 4417 (NY)

Pseudosagedia rhaphidosperma (Müll. Arg.) R. C. Harris – C: Hodkinson 826 (DUKE)

Psilolechia clavulifera (Nyl.) Coppins – M: Buck 53404 (NY)

Psilolechia lucida (Ach.) M. Choisy – M: SNP 133, 270

Psora pseudorussellii Timdal – M: Wieboldt 11158 (VPI)

Psorula rufonigra (Tuck.) Gotth. Schneider – M: Guccion 1077 (NY)

Punctelia appalachensis (Culb.) Krog – C: Luttrell 1802 (US); M: Lendemer 3800 (NY), 3801 (NY), Luttrell 4922 (US)

Punctelia bolliana (Mull. Arg.) Krog – M: Guccion 1078 (NY)

Punctelia borreri (Turner) Krog – M: Culberson 10113 (DUKE), 11282 (DUKE)

Punctelia caseana Lendemer & Hodkinson ined. [= P. subrudecta auct Amer. sensu Lendemer (2004), see Lendemer & Hodkinson (in press)] – C: Reed 64166 (NY); P: Reed 138253 (NY); M: Reed 79194 (NY), Hodkinson 9047 (Hodkinson, pers. herb.), 9267 (Hodkinson, pers. herb.)

Punctelia graminicola (de Lesd.) Egan – M: Luttrell 4866 (US), SNP 21

Punctelia missouriensis G. Wilh. & Ladd – C: Hodkinson 471 (DUKE), 1728 (NY), 2583 (DUKE), Reed 59389 (NY); P: FHM 3707775-7 (DUKE), Ladd 20781 (NY); M: FHM 3708041-3 (DUKE), Lendemer 3842 (NY)

Punctelia reddenda (Stirton) Krog – M: Dey 5643 (DUKE)

Punctelia rudecta (Ach.) Krog – C: Hodkinson 384 (DUKE), 474 (DUKE), 2037 (NY), Reed 61015 (NY); P: Reed 60408 (NY); M: Reed 58342 (NY)

Pycnora praestabilis (Nyl.) Hafellner – M: Hodkinson 9020 (NY)

Pycnothelia papillaria Dufour – C: Hodkinson 85 (DUKE); P: Reed 62981 (NY); M: Harris 12993 (NY)

Pyrenula cruenta (Mont.) Vainio – C: Nearing _1937 (NY)

Pyrenula cuyabensis (Malme) R. C. Harris – C: Hodkinson 1401 (DUKE)

Pyrenula lucifera R. C. Harris – M: Buck 28318 (NY), 28334 (NY)

Pyrenula pseudobufonia (Rehm) R. C. Harris – C: Hodkinson 723 (DUKE), 828 (NY), Reed 61382 (NY); P: Culberson 4718 (DUKE); M: Harris 12878 (NY), 12890 (NY), Buck 20657 (NY), 20687 (NY)

Pyrenula punctella (Nyl.) Trevisan – C: Hodkinson 2226 (DUKE)

Pyrenula subelliptica (Tuck.) R. C. Harris – C: Hodkinson 829 (NY), 883 (DUKE); M: Leeming 12June1892 (NY), Guccion 1079 (NY)

Pyrrhospora varians (Ach.) R. C. Harris – C: Hodkinson 342 (DUKE), 528 (NY), 1415 (DUKE); P: Guccion 1123 (NY); M: Guccion 1080 (NY)

Pyxine sorediata (Ach.) Mont – C: Reed 137638 (NY); P: Hodkinson 5001 (DUKE), 5019 (DUKE), Reed 63059 (NY); M: Reed 67001 (NY)

Pyxine subcinerea Stirton – C: Hodkinson 505 (DUKE), 2102 (NY); P: Guccion 1124 (NY), Hodkinson 10267 (Hodkinson, pers. herb.); M: Guccion 1125 (NY)

Ramalina americana Hale – C: Hodkinson 1793 (DUKE), Anderson 03Sept1934 (NY); M: Reed 106162 (NY)

Ramalina culbersoniorum La Greca – M: Harris 36661 (norbarbatic acid chemotype; NY), Culberson 20585 (divaricatic acid chemotype; DUKE), Culberson 20584 (4-O-methylhypoprotocetraric acid chemotype; DUKE), Culberson 11421 (stenosporic acid chemotype; DUKE)

Ramalina intermedia (Delise ex Nyl.) Nyl. – M: Harris 36720 (NY), Pollard & Maxon 174 (NY)

Ramalina petrina Bowler & Rundel – M: Hermann 15849 (US; Holotype), Culberson 11391 (DUKE), Hale 18110 (US)

Ramalina pollinaria (Westr.) Ach. – M: Harris 10677 (NY)

Ramalina stenospora Müll. Arg. – C: Anderson 03Sept1934 (in a mixed collection labeled as Ramalina americana Hale; NY), Luttrell 1926 (US)

Ramalina willeyi R. Howe – C: Williams 1581 (US)

Ramboldia russula (Ach.) Kalb, Lumbsch & Elix – M: Ihrman 901 (Shen.)

Rhizocarpon cinereovirens (Müll. Arg.) Vainio – M: Buck 36964 (stictic acid chemotype; NY)

Rhizocarpon geographicum (L.) DC. – M: Culberson 11480 (DUKE), SNP 100, 173

Rhizocarpon grande (Flörke ex Flotow) Arnold ? – M: Anderson _June1936 (NY), SNP 68 (sterile)

Rhizocarpon hochstetteri (Körber) Vainio ? – M: SNP 80

Rhizocarpon infernulum (Nyl.) Lynge f. sylvaticum Fryday M: Guccion 105 (NY), Buck 20651 (NY)

Rhizocarpon reductum Th. Fr. – M: Harris 26876 (NY), SNP 63, 81, 97, 113, 120, 145

Rhizocarpon rubescens Th. Fr. – M: Harris 3599 (NY), 26903 (NY), SNP 81, 87, 97, 113, 120, 145

Rhizocarpon subgeminatum Eitner – M: SNP 175

Rhizoplaca subdiscrepans (Nyl.) R. Sant. – M: Hermann 14848 (NY), SNP 95, 174

Rinodina ascociscana Tuck. – M: Buck 28388 (NY), 39140 (NY)

Rinodina destituta (Nyl.) Zahlbr. – P: Reed 113269 (NY); M: Anderson 09June1936 (NY), Harris 12912 (NY)

Rinodina maculans Müll. Arg. – C: Hodkinson 308 (DUKE), 1345 (NY), 1436 (DUKE), 1896 (DUKE), 1897 (DUKE); P: Hodkinson 10248 (Hodkinson, pers. herb.)

Rinodina papillata H. Magn. – C: Hodkinson 1346 (DUKE)

Rinodina subminuta H. Magn. – M: Buck 53381 (NY)

Rinodina tephraspis (Tuck.) Herre – M: Harris 12896 (NY)

Ropalospora chlorantha (Tuck.) S. Ekman – M: Buck 20677 (NY), 53391 (NY)

Ropalospora lugubris (Sommerf.) Poelt – M: SNP 80

Roselliniella sp. ?*– M: Harris 54079 (on Physcia stellaris; NY)

Sarcogyne clavus (DC.) Kremp. – M: SNP 26, 166

Sarcogyne privigna (Ach.) A. Massal. – P: Hodkinson 10208 (DUKE, NY); M: SNP 151, 175

Sarcogyne regularis Körber – M: Ihrman 458-R (Shen.)

Sarcogyne similis H. Magn. – M: Slaughter _1975-76 (EIU)

Sclerococcum sp.* – M: Harris 12989 (on Pertusaria plittiana; NY)

Scoliciosporum chlorococcum (Stenh.) Vězda – M: Buck 36975 (NY), 20667 (NY)

Scoliciosporum pensylvanicum R.C. Harris – M: Guccion 1129 (NY)

Scoliciosporum umbrinum (Ach.) Arnold – M: Anderson 29June1935 (NY), 09July1936 (NY), SNP 75, 175

Segestria leptalea (Durieu & Mont.) R. C. Harris – M: Hodkinson 8979 (NY)

Sphaerellothecium coniodes (Nyl.) Cl. Roux & Diederich* – M: Buck 53348 (on Baeomyces rufus; NY)

Sphinctrina turbinata (Pers.: Fr.) De Not. – M: Harris 26904 (NY), 36671 (NY)

Stereocaulon dactylophyllum Flörke – M: Hale 98 (DUKE)

Stereocaulon glaucescens Tuck. – M: Guccion 1126 (NY), SNP 40, 136, 137, 153, 165

Sticta beauvoisii Delise – M: Buck 44261 (NY), 44262 (NY), Harris 36740 (NY)

Sticta fuliginosa (Hoffm.) Ach. – M: Harris 26994 (NY), Culberson 11333 (DUKE)

Sticta sylvatica (Hudson) Ach. ? – M: Slaughter _1975-76 (EIU)

Stigmidium sp. ?* – M: SNP 268 (on Parmelia omphalodes)

Strigula stigmatella (Ach.) R. C. Harris – M: Buck 53323 (NY)

Strigula submuriformis (R.C. Harris) R.C. Harris – M: Slaughter _1975-76 (EIU)

Teloschistes chrysophthalmus (L.) Th. Fr. – C: Britton 30March1894 (NY); P: Hale 91 (DUKE)

Tephromela atra (Hudson) Hafellner – C: Reed 61667 (NY); M: Anderson 29June1935 (NY), _June1936 (NY)

Thelidium pyrenophorum (Ach.) Mudd – M: Green _1884 (NY), Hodkinson 9312 (NY)

Thelidium zwackhii (Hepp) A. Massal. – M: Hodkinson 9118 (NY)

Thelotrema subtile Tuck. – P: Harris 36641 (NY)

Tomasellia americana (Minks ex Willey) R. C. Harris†‡ – C: Hodkinson 1131 (NY)

Trapelia coarctata (Turner ex Sm. & Sow.) M. Choisy – P: Hodkinson 6124 (DUKE); M: Buck 20690 (NY), Harris 26943 (NY), Hodkinson 9341 (in a mixed collection labeled as Arthonia lapidicola ?; NY)

Trapelia corticola Coppins & P. James – M: Harris 54123 (NY)

Trapelia glebulosa (Sm.) J. R. Laundon – M: Harris 26899 (NY), SNP 63, 73, 145

Trapelia placodioides Coppins & P. James – M: Anderson 29June1935 (NY), Hodkinson 9194 (DUKE)

Trapeliopsis flexuosa (Fr.) Coppins & P. James – C: Hodkinson 2066 (NY), 2195 (DUKE), 2300 (DUKE), 2304 (DUKE); M: Harris 25871 (NY)

Trapeliopsis gelatinosa (Flörke) Coppins & P. James – M: Harris 54124 (NY)

Trapeliopsis granulosa (Hoffm.) Lumbsch – M: Anderson 08June1936 (NY), SNP 105

Trapeliopsis viridescens (Schrader) Coppins & P. James – M: Harris 26965 (NY), 36674 (NY)

Trematosphaeriopsis parmeliana (Jacz.) Elenkin* – M: Hafellner 2809 (on Xanthoparmelia conspersa; Hafellner, pers. herb.)

Tremella parmeliarum Diederich* – M: Hodkinson 9228 (on Parmotrema reticulatum; Hodkinson, pers. herb.), 9235 (on P. reticulatum; Hodkinson, pers. herb.)

'Tremella sp. 7 (Diederich 2007)'* – M: Hodkinson 9019 (on Flavoparmelia caperata; Hodkinson, pers. herb.), 9315 (on F. caperata; Hodkinson, pers. herb.)

Trypethelium virens Tuck. ex E. Michener – C: Hodkinson 1127 (NY), 2416 (DUKE), Reed 59062 (NY); P: Johnson 5255 (NY), 5258 (NY), 5260 (NY); M: Harris 36702 (NY)

Tuckermanella fendleri (Nyl.) Essl. – C: Hodkinson 1126 (DUKE), 1139 (NY), Reed 64643 (NY); P: Culberson 4680 (DUKE); M: Culberson 11447 (DUKE)

Tuckermannopsis americana (Sprengel) Hale – P: Hale 4198 (DUKE); M: Reed 75315 (NY)

Tuckermannopsis ciliaris (Ach.) Gyelnik – C: Reed 57535 (NY); P: Reed 62506 (NY); M: Reed 68921 (NY)

Tuckermannopsis orbata (Nyl.) M. J. Lai – M: Culberson 11389 (NY), Dey 5593 (NY)

Tylothallia biformigera (Leighton) P. James & Kilias – M: Guccion 137 (NY)

Umbilicaria mammulata (Ach.) Tuck. – M: Anderson 10June1936 (NY), Buck 28361 (NY), Harris 36707 (NY), Murrill 24-27Oct1916 (NY)

Umbilicaria muhlenbergii (Ach.) Tuck. – M: Anderson 08June1936 (NY), Small 226 (NY), SNP 68, 174

Usnea amblyoclada (Müll. Arg.) Zahlbr. – M: Reed 66976 (NY), 88984 (NY)

Usnea ceratina Ach. – M: Harris 26948 (NY), 26970 (NY)

Usnea cornuta Körber – M: Dey 5659 (DUKE)

Usnea dasaea Stirton – M: Hodkinson  9203 (NY), 9222 (DUKE), Reed 67623 (NY), 68911 (NY)

Usnea endochrysea Stirton – C: Hodkinson 2019 (originally reported as "Usnea strigosa (Ach.) Eaton" by Hodkinson & Case, 2008; NY), Reed 86824 (NY); M: Reed 1981-107 (NY), Seymour 01Aug1891 (NY)

Usnea fragilescens var. mollis (Vainio) Clerc – M: Dey 5648 (DUKE)

Usnea halei P. Clerc – M: Harris 26879 (NY), Lendemer 4302 (NY), 4303 (NY), SNP 59, Hermann 15847 (DUKE), Culberson 11501 (DUKE)

Usnea lapponica Vainio – M: Seymour 27July-01August1891 (NY), Culberson 11259 (DUKE), 11422 (DUKE), 11436 (DUKE)

Usnea mutabilis Stirton – C: Hodkinson 700 (DUKE), 705 (DUKE), 1838 (NY); M: Hermann 15153 (DUKE)

Usnea pensylvanica Mot. – C: Hodkinson 1328 (NY), 2250 (DUKE), Britton 01April1893 (NY); P: Hale 14611 (DUKE); M: Harris 10652 (NY), Culberson 11375 (DUKE), 11395 (DUKE)

Usnea strigosa (Ach.) Eaton – C: Hodkinson 394 (DUKE), 921 (DUKE), 1452 (DUKE), Reed 139464A (NY), 141968 (NY); P: Harris 36645 (NY); M: Harris 9952 (NY), Leonard 16369 (NY), Leeming 12June1892 (NY)

Usnea subfloridana Stirton – M: Culberson 11501 (DUKE), Harris 54125 (NY)

Usnea subfusca Stirton – M: Allard 9779 (US), Reed 79295 (NY)

Usnea subgracilis Vainio – M: Buck 28354 (NY), Culberson 11396 (DUKE), Guccion 1081 (NY)

Usnea subscabrosa Nyl. ex Mot. – C: Reed 9094 (NY), 128050 (NY); M: Hodkinson 8956 (NY), Reed 67584B (NY)

Usnea trichodea Ach. – C: Reed 46168 (NY), 46162 (diffractaic acid chemotype; NY)

Verrucaria calkinsiana Servít – C: Hodkinson 1293 (DUKE); M: Harris 12871 (NY)

Vouauxiella lichenicola (Lindsay) Petrak & H. Sydow* – M: Slaughter _1975-76 (on Lecanora hybocarpa; filed under Lecanora caesiorubella, site 3; EIU)

Vulpicida viridis (Schwein.) J.-E. Mattsson & M. J. Lai – P: Reed 59650 (NY)

Xanthomendoza fallax (Hepp ex Arnold) Søchting, Kärnefelt & S. Kondr. – C: Hodkinson 1187 (DUKE)

Xanthomendoza weberi (S. Kondr. & Kärnefelt) L. Lindblom – C: Hodkinson 1490 (DUKE), 1539 (NY), 1747 (DUKE); P: Guccion 1082 (NY)

Xanthoparmelia angustiphylla (Gyelnik) Hale – M: SNP 87, 97

Xanthoparmelia conspersa (Ehrh. ex Ach.) Hale – M: Harris 12992 (NY), 36790 (NY), Culberson 11308 (DUKE)

Xanthoparmelia cumberlandia (Gyelnik) Hale – M: Harris 26914 (NY), Buck 20665 (NY)

Xanthoparmelia hypofusca (Gyelnik) Hodkinson & Lendemer ined. [= X. tasmanica auct. Amer., see Hodkinson & Lendemer (in review)] – M: Harris 13005 (NY)

Xanthoparmelia hypomelaena (Hale) Hale – P: Hale 30683 (US)

Xanthoparmelia plittii (Gyelnik) Hale – C: Hodkinson 2444 (DUKE); M: Culberson 11336 (DUKE), Harris 26923 (NY), SNP 19, 20

Xanthoparmelia viriduloumbrina (Gyelnik) Lendemer – M: Harris 36710 (NY)

Xanthoria elegans (Link) Th. Fr. – M: Hale & Poelt _1970 (GZU)

 

Acknowledgements

The authors would like to thank James Lendemer for examining numerous specimens and for providing valuable comments and contributions to this manuscript, Sarah Hodkinson for assistance with collecting, processing, and databasing of specimens, Jolanta Miadlikowska for verifying Peltigera specimens, Phyllis Ihrman (whose cited specimens were all collected in Shenandoah National Park), John G. Guccion, Gary Perlmutter, and Thomas F. Wieboldt for providing specimens representing several new records, and all of those involved with the Forest Health Management Project, the Shenandoah National Park Rock Outcrop Management Project (esp. Wendy Cass), and the 2008 Blomquist Bryological and Lichenological Foray (esp. Fred Huber and Mike Donahue) for helping with crucial data collection. Thanks are also due to Heather M. Wiseman for assisting with field work and Irwin M. Brodo for support and training through his lichen identification class at the Humboldt Field Research Institute. Gratitude is extended to Connie Robertson and Molly McMullen for their assistance in the preparation, storage, and databasing of specimens. The authors also thank Suzanne Joneson, Cécile Gueidan, Tami McDonald, and François Lutzoni for examining specimens, Gregory McKee and Rusty Russell for assistance with US collections, and Chicita Culberson for instruction in thin layer chromatography. Thanks must be extended to the numerous collectors that have generously deposited their specimens in various herbaria, making them accessible for examination as part of this project. Support for this research was provided by a grant to the College of William and Mary from the Howard Hughes Medical Institute Undergraduate Science Education Program. Funding was also provided by the William and Mary Charles Center and an Adkins Arboretum Botanical Research Award to BPH.

 

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________________________________________________________________________

Last Updated: 05 January 2010

Maintained by: Brendan Hodkinson

If you have any records worth examining or see any potential errors please contact me at bph8@duke.edu. Thanks for visiting the site!
-Brendan